Understanding Mixture-of-Experts Language Models Beyond Expert Boundaries

In Mixture-of-Experts (MoE) models, the nature of expert specialization remains controversial. Reports of domain-associated expert preferences contrast with analyses attributing routing primarily to token identity or syntactic patterns. To clarify these differing accounts, we look beyond expert-level routing to investigate the neuron-level organization of domain-specialized computation using Routing-Aware Magnitude Profiling (RAMP), a measure developed for this analysis. Using mixed-domain calibration data, we identify domain-specialized neuron sets with 58-74% overlap across multi-source samples of the same domain, compared with 15-24% across domains. To understand how this organization evolves during training, we trace neuron importance throughout pre-training and post-training across three fully open MoE families. Neuron importance continues to reorganize between pre-training checkpo…